In the early 19th century, the Bennet family lives at their Longbourn estate, situated near the small town of Meryton in Hertfordshire, England. Mrs Bennet's greatest desire is to marry off her five daughters to secure their futures. A regiment of militia is encamped outside the town, and the younger Bennet daughters flirt with the officers, to the great embarrassment of their elder sisters.
The arrival of Mr Bingley, a rich bachelor who rents the neighbouring Netherfield estate, gives her hope that one of her daughters might contract a marriage to their advantage. At a ball, the family is introduced to the Netherfield party: Mr Bingley; his sisters and brother-in-law; and his closest friend Mr Darcy. Mr Bingley appears interested in Jane, the eldest Bennet daughter, while Mr Darcy, reputed to be twice as wealthy as Mr Bingley, is haughty and aloof, alienating the village. He declines to dance with Elizabeth, the second-eldest Bennet daughter, as she is "not handsome enough." Although she jokes about it, Elizabeth is deeply offended. Despite this first impression, Mr Darcy secretly finds himself drawn to Elizabeth as they continue to encounter each other at social events, appreciating her wit and frankness.
Mr Collins, a recently ordained clergyman and (as a distant cousin) the heir to the Longbourn estate, which is entailed away from the daughters, visits the Bennet family. He intends to marry one of the Bennet girls on the advice of his patroness Lady Catherine de Bourgh, who is also Mr Darcy's aunt. He decides to pursue Elizabeth. The Bennet family meets the charming militia officer George Wickham, who tells Elizabeth in confidence that Mr Darcy had treated him badly in the past. Elizabeth's prejudice against Mr Darcy and admiration for Wickham lead her to believe him. Elizabeth rejects Mr Collins's marriage proposal, and Mr Collins subsequently proposes to Charlotte Lucas, a friend of Elizabeth's, and is accepted. Convinced that Jane is an unsuitable match for his friend, Mr Darcy prompts Mr Bingley to leave for London and, supported by the two sisters, persuades him not to return to Netherfield. Jane is heartbroken, while Elizabeth's hatred for Mr Darcy grows, as she correctly suspects he was responsible for Mr Bingley's departure.
In the spring, Elizabeth visits Charlotte and Mr Collins in Kent. Elizabeth and her hosts are invited to Rosings Park, Lady Catherine's home, where Mr Darcy and his cousin Colonel Fitzwilliam are also visiting. Fitzwilliam unknowingly confirms Elizabeth's suspicion that Mr Darcy had intentionally separated Bingley and Jane. Mr Darcy proposes to Elizabeth, declaring his love for her despite her low social connections. She is shocked and rejects him angrily, saying that she could never love a man who caused her sister such unhappiness; she further accuses him of treating Wickham unjustly. Darcy admits to separating Bingley and Jane but dismisses the accusation regarding Wickham. The next day, Mr Darcy gives Elizabeth a letter explaining his history with Wickham, which contradicts Wickham's earlier assertions about their relationship and paints Wickham as profligate and predatory to young women, including Darcy's teenage sister. Darcy explains but does not apologize for his efforts to separate Bingley and Jane, whom he believed indifferent to his friend. Though Elizabeth knew her sister was deeply in love, the letter convinces her that Darcy is a good, though proud, man and she had been unfair to him about Wickham.