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An actor (masculine/gender-neutral), or actress (feminine), is a person who portrays a character in a production. The actor performs "in the flesh" in the traditional medium of the theatre or in modern media such as film, radio, and television. The analogous Greek term is ὑποκριτής (hupokritḗs), literally "one who answers". The actor's interpretation of a role—the art of acting pertains to the role played, whether based on a real person or fictional character. This can also be considered an "actor's role", which was called this due to scrolls being used in the theaters. Interpretation occurs even when the actor is "playing themselves", as in some forms of experimental performance art.

Formerly, in ancient Greece and the medieval world, and in England at the time of William Shakespeare, only men could become actors, and women's roles were generally played by men or boys. While Ancient Rome did allow female stage performers, only a small minority of them were given speaking parts. The commedia dell'arte of Italy, however, allowed professional women to perform early on; Lucrezia Di Siena, whose name is on a contract of actors from 10 October 1564, has been referred to as the first Italian actress known by name, with Vincenza Armani and Barbara Flaminia as the first primadonnas and the first well-documented actresses in Italy (and in Europe). After the English Restoration of 1660, women began to appear onstage in England. In modern times, particularly in pantomime and some operas, women occasionally play the roles of boys or young men.

While the word actor has existed for much of the history of English language, as to mean "one who does something", it was not used to refer to one who performs in theatre until the 16th century.

The profession of acting possesses a significant amount of terminology, some of which is historically or contemporaneously contentious. This includes the term actress and less common player.

According to the OED, the earliest recorded occurrence of the term "actress" in reference to stage performance came in 1608 in the anonymous play Famelie of Love.

Seventeenth-century English writers sometimes used the forms "actoress" or "actoresse" to refer to a female appearing on stage.

In the 19th century, many people viewed women in acting negatively, as actresses were often courtesans; females on stage had become associated with promiscuity. Despite such prejudices, the 19th century also saw early female stage "stars", most notably Sarah Bernhardt.

With regard to the cinema of the United States, the gender-neutral term "player" was common in the silent film era and the early days of the Motion Picture Production Code, but in the 2000s in a film context, it is generally deemed archaic. However, "player" remains in use in the theatre, often incorporated into the name of a theatre group or company, such as the American Players, the East West Players, etc. Also, actors in improvisational theatre may be referred to as "players".

The first recorded case of a performing actor occurred in 534 BC (though the changes in the calendar over the years make it hard to determine exactly) when the Greek performer Thespis stepped onto the stage at the Theatre Dionysus to become the first known person to speak words as a character in a play or story. Before Thespis' act, Grecian stories were only expressed in song, dance, and in third person narrative. In honor of Thespis, actors are commonly called Thespians. The exclusively male actors in the theatre of ancient Greece performed in three types of drama: tragedy, comedy, and the satyr play. This developed and expanded considerably under the Romans. The theatre of ancient Rome was a thriving and diverse art form, ranging from festival performances of street theatre, nude dancing, and acrobatics, to the staging of situation comedies, to high-style, verbally elaborate tragedies.

As the Western Roman Empire fell into decay through the 4th and 5th centuries, the seat of Roman power was moved eastward to Constantinople. Records show that mime, pantomime, scenes or recitations from tragedies and comedies, dances, and other entertainments were very popular. From the 5th century, Western Europe was plunged into a period of general disorder. Small nomadic bands of actors traveled around Europe throughout the period, performing wherever they could find an audience; there is n